1993-1997 BA Natural Sciences, Emmanuel College, Cambridge University, UK
1997-2002 PhD with Julian Downward and Gerard Evan (now UCSF) at Imperial Cancer Research Fund/Cancer Research UK
2002-2007 Postdoc with Michael Cole at Dept. of Molecular Biology, Princeton University, USA and Dept. of Pharmacology and Toxicology, Dartmouth College, USA
2007- Principal Investigator at Dundee University, College of Life Sciences, Division of Cell Biology and Immunology

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